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Culture media were refreshed daily. Cells were maintained at 37 C and 5% CO2 in a humidified incubator and regularly tested for mycoplasma contamination.\n\nGeneration of Cortical Brain Organoids: Cortical brain organoids were generated from hiPSCs as previously described. Briefly, hiPSCs were dissociated into single cells using TrypLE, and 5.0 \u00d7 10^5 cells were seeded per well in ultra-low attachment U-bottom 96-well plates (Corning) to form uniform aggregates in mTeSR Plus medium (STEMCELL Technologies) supplemented with 50 uM Y-27632 (BioGems). On day 2, two-thirds of the medium was replaced every other day with fresh mTeSR Plus medium lacking Y-27632. 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